Mweene - delighted to see a John Mikel Obi missed spot-kick in the first half - coolly planted the ball into the top right corner after Nigeria defender Ogenyi Onazi was harshly adjudged to have fouled Emmanuel Mayuka.
Nigeria had led through a 57th-minute strike from the impressive Emmanuel Emenike, with the Super Eagles on balance the better side but Zambia having also spurned some great chances before finally drawing level.
The result means defending champions Zambia and two-times winners Nigeria have drawn their opening two matches at these finals.
Nigeria will travel to Rustenburg to face Ethiopia, Zambia will stay in Mbombela to play group leaders Burkina Faso in Tuesday's final group matches.
Video highlight: Zambia 1-1 Nigeria http://www.footytube.com/video/zambia-nigeria-jan25-154580?ref=lchan_vidgrid
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